ATHENS, Greece -- Five suspects implicated in the killing of a University of California, Berkeley professor appeared in court in Athens on Thursday, including his ex-wife, who has denied all charges.

Greek officials say the broad-daylight attack came one day after a custody hearing over the victim's 10-year-old twins.

Greek police have arrested five people over the killing of a University of California at Berkeley professor who was shot dead in an Athens suburb earlier in July, police said.